Based on the same-name Game Cube and Play Station game, situation at the end of Tales of Symphonia, The Animation. Two worlds exist, both unaware of the other, as two ends of an hourglass where sand is mana. In times, the Angels grant their blessing to a Chosen One who must travels to awaken the Spirits guarding mana: that is the world regeneration. Sylvarant is the homeland of the Chosen Colette and her friends, Lloyd, Genis and Raine. When they reached the final stage of their quest, the true nature of the world regeneration was revealed: the Chosen must sacrifice herself and become a vessel for the Goddess Martel. Refusing to let Colette face this fate, Lloyd opposed the Angels, despite the betrayal of Kratos, and they run away to the other world, Thethe'alla. The balance and the secrecy between the two worlds begin to disappear, and Lloyd and his friends must now pursue their quest in Thethe’alla to save Colette and prevent both worlds to crumble.

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